Zhang L, Qin W, Ishii S, Stahl DA, Winkler MK. Oxygen dependent nitric oxide (NO) and nitrous oxide (N2O) dynamics during aerobic ammonia oxidation. Journal of Microbiological Methods. 2026 Jul 9:107628.

Flinkstrom Z, Hunt KA, Abrahamson B, Harvell P, Li X, Wilson J, Cooper ZS, Valenzuela JJ, Baliga NS, Stahl DA, Qin W, Winkler MH. Comparative physiological and genomic characterization of a novel Nitrobacter vulgaris strain from a nitrate-contaminated subsurface. Appl Environ Microbiol 0:e00130-26.

Yang X, Zhao B, Feng K, Wang J, Liu M, Peng X, He Q, Lu Y, Waseem H, Wang S, Winkler MKH. 2026. Microbial synthesis structures organic compound composition in anaerobic digestion. The ISME Journal, p.wrag036.

Wang PH, Saingam P, Rasyid R, Godfrey BJ, Himmelfarb J, Winkler MKH. 2026. Indole toxicity on removal of uremic toxin p-cresol, in-vitro study of Thauera aminoaromatica S2. bioRxiv, pp.2026-01.

Sen O, Wang PH, Saingam P, Godfrey BJ, Himmelfarb J, Xiong Y, Pan C, Winkler MKH. 2026. Biochemical mechanism of p-cresol removal by Thauera aminoaromatica S2. bioRxiv, pp.2026-01.

Godfrey BJ, Wang PH, Saingam P, Tju H, Winkler MKH. 2026. Development of a Stabilized Alginate-based Hydrogel for Oral Delivery of Encapsulated Live Cultures and Allowing their Intact Passage Through the Digestive System. bioRxiv, pp.2026-01.

Hong S, Winkler MK, Wang ZW, Dhanasekar A, Goel R. 2026. Unveiling Metabolic Insights and Niche Differentiation of Microbial Communities in EBPR-Anammox Reactor through Integration of Long-Read Metagenomics and Metatranscriptomics. Environmental Science & Technology.

Zhou L, Dong N, Ou P, Shen Y, Jian Y, Winkler MK, Rittmann BE. 2025. Granule Deterioration Triggers Increased Dissimilatory Nitrate Reduction to Ammonium in Anammox Granular Sludge. ACS ES&T Engineering, 5(8), pp.1985-1994.

Bloom PP, Garrett WS, Penniston KL, Winkler MKH, Hazen SL, Agudelo J, Suryavanshi M, Babiker A, Dodd D, Fischbach MA, Huang KC. 2025. Microbiota and kidney disease: the road ahead. Nature Reviews Nephrology, pp.1-15.

Mohammadi R, Eshaq G, Winkler MK, Pihlajamki A. 2025. Recent Development of Novel Magnetic Biochar Aluminum Cross-Linked Composite Beads for Preferential Phosphate Separation from Phosphate-Rich Effluents. Available at SSRN 4555358.

Saingam P, Rasyid R, Abrahamson B, Lie TJ, Godfrey B, Himmelfarb J, Winkler MKH. 2025. Towards an effective delivery system of a microbial sink of the uremic toxin, p-cresol; an in vitro study with Thauera aminoaromatica S2. Frontiers in Microbiology, 16, p.1577556.

Davis R, Mafune K, Winkler MKH. 2025. Biodegradable Hydrogels and Microbial Consortia as a Treatment for Soil Dysbiosis. Frontiers in Microbiology, 16, p.1565940.

Kwon H, Li B, Xu M, Wang Q, Maqbool T, Lu H, Winkler MK, Jiang D. 2025. Minimizing byproduct formation in bioelectrochemical denitrification with annamox bacteria. Journal of Hazardous Materials, p.138110.

Mafune KK, Winkler MK. 2024. The expansion of fungal organisms in environmental biotechnology. Current Opinion in Biotechnology, 90, p.103217.

Brownlie WJ, Aziz T, Sutton MA, Shenk A, Winkler MKH, Robinson G, Adhya TK, Amon B, Awan MI, Bittman S, Bleeker A. 2024. Nitrogen Mitigation: INMS Guidance Document on Measures for Sustainable Nitrogen Management.

Wang J, Sun Y, Zhang X, Khunjar W, Bo L, Winkler MK, Goel R, Wang ZW. 2024. Carbon-Efficient Nutrients Removal from Real Municipal Wastewater Under Conditions of Highly Variable Influent Quality and Low Temperature. Available at SSRN 4861845.

Mafune KK, Kasson MT, Winkler MKH. 2024. Building blocks toward sustainable biofertilizers: variation in arbuscular mycorrhizal spore germination when immobilized with diazotrophic bacteria in biodegradable hydrogel beads. Journal of Applied Microbiology, 135(7).

Johnston JT, Nguyen Quoc B, Abrahamson B, Candry P, Ramon C, Cash KJ, Saccomano SC, Samo TJ, Ye C, Weber PK, Winkler MKH, Mayali X. 2024. Increasing aggregate size reduces single-cell organic carbon incorporation by hydrogel-embedded wetland microbes. ISME Communications, ycae086.

Wang D, Candry P, Hunt K, Flinkstrom Z, Winkler MKH, Stahl DA, Pan C. 2024. Metaproteomics-informed stoichiometric modeling reveals the responses of wetland microbial communities to oxygen and sulfate exposure. npj Biofilms Microbiomes 10, 55.

Flinkstrom Z, Bryson S, Candry P, Winkler MKH. 2024. Metagenomic clustering links specific metabolic functions to globally relevant ecosystems. mSystems, pp.e00573-24.

Candry P, Godfrey BJ, Winkler MKH. 2024. Microbe-cellulose hydrogels as a model system for particulate carbon degradation in soil aggregates. ISME Communications, 4(1), p.ycae068.

Saingam P, Jain T, Woicik A, Li B, Candry P, Redcorn R, Wang S, Himmelfarb J, Bryan A, Winkler MKH. 2024. Integrating Socio-Economic Vulnerability Factors Improves Neighborhood-Scale Wastewater-Based Epidemiology for Public Health Applications. Water Research.

Qin W, Wei SP, Zheng Y, Abrahamson B, Flinkstrom Z, Hunt KA, Stahl DA, Winkler MKH, et al. 2024. Ammonia-oxidizing bacteria and archaea exhibit differential nitrogen source preferences. Nat Microbiol.

Nguyen Quoc B, Cavanaugh SK, Hunt KA, Bryson SJ, Winkler MKH. 2024. Impact of aerobic granular sludge sizes and dissolved oxygen concentration on greenhouse gas N2O emission. Water Research, p.121479.

Nguyen Quoc B, Peng B, De Clippeleir H, Winkler MKH. 2024. Case study: Bioaugmenting the comammox dominated biomass from B-stage to enhance nitrification in A-stage at Blue Plains AWWTP. Water Environment Research, 96(3).

Zheng Y, Wang B, Gao P, Yang Y, Xu B, Su X, Ning D, Tao Q, Li Q, Winkler MKH, Zhao F, Wang D. 2024. Novel order-level lineage of ammonia-oxidizing archaea widespread in marine and terrestrial environments. The ISME Journal, 18(1), p.wrad002.

Horn EK, Joyce A, Chowdhury RB, Caputo S, Jacobs B, Winkler MKH, Proksch G. 2023. Translating Environmental Potential to Economic Reality: Assessment of Commercial Aquaponics through Sustainability Transitions Theory. Circular Economy and Sustainability, pp.1-32.

Candry P, Flinkstrom Z, Winkler MKH. 2023. Wetlands harbor lactic acid-driven chain elongators. Microbiology Spectrum, pp.e02105-23.

Candry P, Chadwick GL, Caravajal-Arroyo JM, Lacoere T, Winkler MKH, Ganigué R, Orphan VJ, Rabaey K. 2023. Trophic interactions shape the spatial organization of medium-chain carboxylic acid producing granular biofilm communities. ISME J.

Godfrey B, Li B, Gottshall E, Bryson S, Abrahamson B, Winkler MKH. 2023. Co-immobilization of AOA Strains with Anammox Bacteria in Three Different Synthetic Bio-granules Maintained under Two Substrate-level Conditions. Chemosphere, 342, 140192.

Xing B, Graham NJD, Zhao B, Li X, Tang Y, Kappler A, Dong H, Winkler MKH, Yu W. 2023. Goethite Formed in the Periplasmic Space of Pseudomonas sp. JM-7 during Fe Cycling Enhances Its Denitrification in Water. Environ. Sci. Technol.

Kim Y, Flinkstrom Z, Candry P, Winkler MKH, Myung J. 2023. Resource availability governs polyhydroxyalkanoate (PHA) accumulation and diversity of methanotrophic enrichments from wetlands. Frontiers in Bioengineering and Biotechnology, 11, p.1210392.

Candry P, Abrahamson B, Stahl DA, Winkler MKH. 2023. Microbially mediated climate feedbacks from wetland ecosystems. Global Change Biology.

Li B, Godfrey BJ, RedCorn R, Candry P, Abrahamson B, Wang Z, Goel R, Winkler MKH. 2023. Mainstream Nitrogen Removal from Low Temperature and Low Ammonium Strength Municipal Wastewater using Hydrogel-encapsulated Comammox and Anammox. Water Research, 242, p.120303.

Li B, Godfrey BJ, RedCorn R, Wang Z, Goel R, Winkler MKH. 2023. Simultaneous anaerobic carbon and nitrogen removal from primary municipal wastewater with hydrogel encapsulated anaerobic digestion sludge and AOA-anammox coated hollow fiber membrane. Science of The Total Environment, 883, p.163696.

Daigger GT, Kuo J, Derlon N, Houweling D, Jimenez JA, Johnson BR, McQuarrie JP, Murthy S, Regmi P, Roche C, Sturm B, Wett B, Winkler MKH, Boltz JP. 2023. Biological and Physical Selectors for Mobile Biofilms, Aerobic Granules, and Densified-Biological Flocs in Continuously Flowing Wastewater Treatment Processes: A State-of-the-Art Review. Water Research, p.120245.

Wang D, Hunt KA, Candry P, Tao X, Wofford NQ, Zhou J, McInerney MJ, Stahl DA, Tanner RS, Zhou A, Winkler MKH, Pan C. 2023. Cross-Feedings, Competition, and Positive and Negative Synergies in a Four-Species Synthetic Community for Anaerobic Degradation of Cellulose to Methane. mBio.

Hong S, Winkler MKH, Wang Z, Goel R. 2023. Integration of EBPR with mainstream anammox process to treat real municipal wastewater: process performance and microbiology. Water Research, p.119758.

Saingam P, Li B, Nguyen Quoc B, Jain T, Bryan A, Winkler MKH. 2023. Wastewater surveillance of SARS-CoV-2 at intra-city level demonstrated high resolution in tracking COVID-19 and calibration using chemical indicators. Science of The Total Environment, 866:161467.

Gottshall EY, Godfrey B, Li B, Abrahamson B, Winkler MKH, Qin W. 2022. Photoinhibition of Comammox reaction in Nitrospira inopinata in a dose and wavelength-dependent manner. Frontiers in Microbiology, p.4964.

Candry P, Godfrey BJ, Wang Z, Sabba F, Dieppa E, Fudge J, Balogun O, Wells G, Winkler MKH. 2022. Tailoring polyvinyl alcohol-sodium alginate (PVA-SA) hydrogel beads by controlling crosslinking pH and time. Sci Rep 12, 20822.

Xie T, Liu X, Xu Y, Bryson S, Zhao L, Huang K, Huang S, Li X, Yang Q, Dong H, Winkler MKH. 2022. Coupling methanotrophic denitrification to anammox in a moving bed biofilm reactor for nitrogen removal under hypoxic conditions. Science of The Total Environment, 856, p.158795.

Zhang W, Wei SP, Winkler MKH, Mueller AV. 2022. Design of a Soft Sensor for Monitoring Phosphorous Uptake in an EBPR Process. ACS ES&T Engineering.

Cavanaugh SK, Nguyen Quoc B, Jacobson E, Bucher R, Sukapanpotharam P, Winkler MKH. 2022. Impact of nitrite and oxygen on nitrous oxide emissions from a granular sludge sequencing batch reactor. Chemosphere, p.136378.

Nguyen Quoc B, Saingam P, RedCorn R, Carter JA, Jain T, Candry P, Gattuso M, Huang MW, Greninger AL, Meschke JS, Bryan A, Winkler MKH. 2022. Case Study: Impact of Diurnal Variations and Stormwater Dilution on SARS-CoV-2 RNA Signal Intensity at Neighborhood Scale Wastewater Pumping Stations. ACS ES&T Water.

Bryson SJ, Hunt KA, Stahl DA, Winkler MKH. 2022. Metagenomic insights into competition between denitrification and DNRA within one-stage and two-stage partial-nitritation Anammox bioreactor configurations. Frontiers in Microbiology, p.1402.

Winkler MKH, Van Loosdrecht MCM. 2022. Intensifying existing urban wastewater. Science, Vol 375, Issue 6579, pp. 377-378.

RedCorn R, Jake JL, Gottshall E, Stahl DA, Winkler MKH. 2021. Light-Weight Oxygen Supply for Portable Biological Nitrogen Removal from Urine and Sweat. Chemical Engineering Journal Advances, p.100235.

Pelivano B, Bryson S, Hunt KA, Denecke M, Stahl DA, Winkler MKH. 2021. Application of pyritic sludge with an anaerobic granule consortium for nitrate removal in low carbon systems. Water Research, p.117933.

Landreau M, You H, Stahl DA, Winkler MKH. 2021. Immobilization of active ammonia-oxidizing archaea in hydrogel beads. npj Clean Water 4, 43.

Wei SP, Stensel HD, Ziels RM, Herrera S, Lee PH, Winkler MKH. 2021. Partitioning of Nutrient Removal Contribution Between Granules and Flocs in a Hybrid Granular Activated Sludge System. Water Research, 117514.

Gottshall EY, Bryson SJ, Cogert KI, Landreau M, Sedlacek CJ, Stahl DA, Daims H, Winkler MKH. 2021. Sustained nitrogen loss in a symbiotic association of Comammox Nitrospira and Anammox bacteria. Water Research, p.117426.

Yin J, Hunt KA, Xie T, Quoc BN, Tran K, Stahl DA, Winkler MKH. 2021. Pairing denitrifying phosphorus accumulating organisms with anaerobic ammonium oxidizing bacteria for simultaneous N and P removal. Science of The Total Environment, p.147521.

Nguyen Quoc B, Armenta M, Carter J, Bucher R, Sukapanpotharam P, Stahl DA, Stensel HD, Winkler MKH. 2021. An investigation into the optimal granular sludge size for simultaneous nitrogen and phosphate removal. Water Research.

Wei SP, Nguyen Quoc B, Shapiro M, Chang PH, Jason C, Winkler MKH. 2020. Application of Aerobic Kenaf Granules for Biological Nutrient Removal in a Full-scale Continuous Flow Activated Sludge System. Chemosphere, p.129522.

Jia M, Winkler MKH, Volcke E. 2020. Elucidating the competition between heterotrophic denitrification and DNRA using the resource-ratio theory. Environmental Science & Technology, 54(21), pp.13953-13962.

Nguyen Quoc B, Wei S, Armenta M, Bucher R, Sukapanpotharam P, Stahl DA, Stensel HD, Winkler MKH. 2020. Aerobic granular sludge: Impact of size distribution on nitrification capacity. Water Research, p.116445.

Garg A, Stensel HD, Bucher B, Sukapanpotharam P, Winkler MK. 2020. Effect of waste activated sludge pretreatment methods to mitigate Gordonia foaming potential in anaerobic digestion. Water and Environment Journal.

Weissbrodt DG, Winkler MKH, Wells GF. 2020. Responsible Science, Engineering and Education for Water Resource Recovery and Circularity. Environ. Sci.: Water Res. Technol., 6, 1952-1966.

Qin W, Zheng Y, Zhao F, Wang Y, Urakawa H, Martens-Habbena W, Liu H, Huang X, Zhang X, Nakagawa T, Mende DR, Bollmann A, Wang B, Zhang Y, Amin SA, Nielsen JL, Mori K, Takahashi R, Armbrust EV, Winkler MKH, DeLong DF, Li M, Lee PH, Zhou J, Zhang C, Zhang T, Stahl DA, Ingalls AE. 2020. Alternative strategies of nutrient acquisition and energy conservation map to the biogeography of marine ammonia-oxidizing archaea. The ISME Journal, pp.1-5.

Landreau M, Byson SJ, You H, Stahl DA, Winkler MKH. 2020. Effective nitrogen removal from ammonium-depleted wastewater by partial nitritation and anammox immobilized in granular and thin layer gel carriers. Water Research, p.116078.

Wei SP, Stensel HD, Nguyen Quoc B, Stahl DA, Huang X, Lee PH, Winkler MKH. 2020. Flocs in disguise? High granule abundance found in continuous-flow activated sludge treatment plants. Water Research.

Nishiguchi DK, Winkler MKH. 2020. Correlating Sludge Constituents with Digester Foaming Risk Using Sludge Foam Potential and Rheology. Water Science and Technology.

Yu Z, Pesesky M, Zhang L, Huang J, Winkler MKH, Chistoserdova L. 2020. A Complex Interplay between Nitric Oxide, Quorum Sensing, and the Unique Secondary Metabolite Tundrenone Constitutes the Hypoxia Response in Methylobacter. mSystems, 5(1).

Cogert K, Ziels R, Winkler MKH. 2019. Reducing Cost and Environmental Impact of Wastewater Treatment with Denitrifying Methanotrophs, Anammox, and Mainstream Anaerobic Treatment. Environ. Sci. Technol.

Pelivanoski B, Detmann B, Ooms K, Winkler MKH, Vasyukova E, Denecke M. 2019. Design of a 1,000 L pilot-scale airlift bioreactor for nitrification with application of a three-phase hydrodynamic mathematical model and prediction of a low liquid circulation velocity. Chemical Engineering Research and Design.

Straka L, Summers A, Stahl DA, Winkler MKH. 2019. Kinetic Implication of moving warm side-stream Anaerobic Ammonium Oxidizing Bacteria to cold mainstream wastewater. Bioresource Biotechnology.

Straka L, Meinhardt KA, Bollmann A, Stahl DA, Winkler MKH. 2019. Affinity informs environmental cooperation between ammonia-oxidizing archaea (AOA) and anaerobic ammonia-oxidizing (Anammox) bacteria. The ISME Journal.

Winkler MKH, Straka L. 2019. New directions in biological nitrogen removal and recovery from wastewater. Current Opinion in Biotechnology, 57, pp.50-55.

Vuono DC, Read RW, Hemp J, Sullivan BW, Arnone JA, Neveux I, Blank B, Staub C, Loney E, Miceli D, Winkler MKH, Chakraborty R, Stahl DA, Grzymski J. 2018. Resource limitation modulates the fate of dissimilated nitrogen in a dual-pathway Actinobacterium. Front. Microbiol.

Snauffer A, Chauhan U, Cogert K, Winkler MKH, Mueller A. 2018. Data Fusion for Environmental Process Control: Maximizing Useful Information Recovery under Data Limited Constraints. IEEE Sensors Letters.

Wei SP, van Rossum F, van de Pol GJ, Winkler MKH. 2018. Recovery of phosphorus and nitrogen from human urine by struvite precipitation, air stripping and acid scrubbing: A pilot study. Chemosphere.

Xie T, Yang Q, Winkler MKH, Wang D, Zhong Y, An H, Chen F, Yao F, Wang X, Wu J, Li X. 2018. Perchlorate bioreduction linked to methane oxidation in a membrane biofilm reactor: Performance and microbial community structure. Journal of Hazardous Materials.

Jia M, Castro-Barros CM, Winkler MKH, Volcke EIP. 2018. Effect of organic matter on the performance and N2O emission of a granular sludge anammox reactor. Environmental Science: Water Research & Technology, 4(7), pp.1035-1046.

Figdore BA, Stensel D, Winkler MKH. 2018. Bioaugmentation of sidestream nitrifying-denitrifying phosphorus-accumulating granules in a low-SRT activated sludge system at low temperature. Water Research.

Figdore BA, Stensel DH, Winkler MKH. 2018. Comparison of different aerobic granular sludge types for activated sludge nitrification bioaugmentation potential. Bioresource Technology: 251: 189-196.

Winkler MKH, Meunier C, Henriet O, Mahillon J, Suárez-Ojeda ME, Del Moro G, De Sanctis M, Di Iaconi C, Weissbrodt DG. 2017. An integrative review of granular sludge for the biological removal of nutrients and recalcitrant organic matter from wastewater. Chemical Engineering Journal.

Figdore BA, Winkler MKH, Stensel D. 2017. Bioaugmentation with Nitrifying Granules in Low-SRT Flocculent Activated Sludge at Low Temperature. Water Environment Research.

Figdore BA, Stensel DH, Winkler MKH, Neethling JB. 2017. Aerobic Granular Sludge for Biological Nutrient Removal. WHITE PAPER, Water Environment Reuse Foundation.

Castro-Barros CM, Ho LT, Winkler MKH, Volcke EIP. 2017. Integration of Methane Removal in Aerobic Anammox-Based Granular Sludge Reactors. Environmental Technology:1-63.

Winkler MKH, Boets P, Hahne B, Goethals P, Volcke EIP. 2017. Effect of the dilution rate on microbial competition: r-strategist can win over k-strategist at low substrate concentration. PLoS ONE 12(3).

Castro-Barros CM, Mingsheng J, van Loosdrecht MCM, Volcke EIP, Winkler MKH. 2017. Evaluating the potential for dissimilatory nitrate reduction by anammox bacteria for municipal wastewater treatment. Bioresource Technology, 233, 363-372.

Winkler MKH, Kröber E, Mohn W, Koch F, Frigon D. 2016. Comparison of microbial populations and foaming dynamics in conventional versus membrane enhanced biological phosphorous removal system. Water and Environment Journal.

Vannecke T, Nicolas B, Winkler MKH, Gaelle S, Steyer J, Volcke EPI. 2016. Influence of process dynamics on the microbial diversity in a nitrifying biofilm reactor. Biotechnology and Bioengineering, 13(9), 1962-1974.

Winkler MKH, Le QH, Volcke EPI. 2015. Influence of partial denitrification and mixotrophic growth of NOB on microbial distribution in aerobic granular sludge reactor. Environmental Science and Technology, 49:11003-11010.

Winkler MKH, Ettwig KF, Vannecke TPW, Stultiens K, Bogdan A, Kartal B, Volcke EPI. 2015. Modelling simultaneous anaerobic methane and ammonium removal in a granular sludge reactor. Water Research, 73:323-331.

Winkler MKH, Bennenbroek MH, Horstink FH, van Loosdrecht MCM, van de Pol GJ. 2013. The biodrying concept: An Innovative Technology creating energy from sewage sludge. Bioresource Technology, 147:124-129.

Zhang Z, Winkler MKH, Wu Z, Lu H, Czapar GF, Wang H, Zheng J. 2013. A Full-scale Housefly Larva (Musca Domestica) Bioconversion for Value-added Swine Manure Stabilization. Waste Resources, 31(2):223-31.

Winkler MKH, van Rossum F, Wicherink B. 2013. Approaches to Urban Mining Recovery of Ammonium and Phosphate from Human Urine. G.I.T. Laboratory Journal.

Winkler MKH, Yang J, Kleerebezem R, Plaza E, Trela, Hultman B, van Loosdrecht MCM. 2012. Nitrate reduction by organotrophic anammox bacteria in a partial nitrifying granular sludge and a moving bed biofilm reactor. Bioresource Technology, 114:217-23.

Winkler MKH, Bassin JP, Kleerebezem R, van der Lans RGJM, van Loosdrecht CM. 2012. Temperature and salt effects on settling velocity in granular sludge technology. Water Research, 46(12):3897-902.

Winkler MKH, Bassin JP, Kleerebezem R, Sorokin D, van Loosdrecht MCM. 2012. Unravelling the reasons for disproportion in the ratio of AOB and NOB in aerobic granular sludge. Applied Environmental Biotechnology, 94(6):1657-66.

Winkler MKH, Kleerebezem R, van Loosdrecht MCM. 2012. Integration of anammox into the aerobic granular sludge process for mainstream wastewater treatment at ambient temperatures. Water Research, 46(1):136-44.

Bassin JP, Winkler MKH, Kleerebezem R, van Loosdrecht MCM. 2012. Improved phosphate removal by selective sludge discharge in aerobic granular sludge reactors. Biotechnology and Bioengineering, 45(11):3291-9.

Winkler MKH, Kleerebezem R, Khunjar W, de Bruin B, van Loosdrecht MCM. 2012. Evaluating the solid retention time of bacteria in flocculent and granular sludge. Water Research, 46(16):4973-80.

Winkler MKH, Kleerebezem R, Strous M, Chandran K, van Loosdrecht MCM. 2012. Factors influencing granular density of aerobic granular sludge. Applied Environmental Biotechnology, (16):7459-68.

Kim YM, Winkler MKH, van Loosdrecht MCM, Chandran K. 2012. The effect of inorganic carbon limitation on nitrite oxidizing bacteria. Proceedings of the Water Environment Research, WEFTEC 2012: Session 21 through Session 30, pp.1968-1974(7).

Winkler MKH, Kleerebezem R, Verhijen P, van Loosdrecht MCM. 2012. Microbial diversity differences within aerobic granular sludge and activated sludge flocs. Applied Environmental Biotechnology, (16):7447-58.

Winkler MKH, Bassin P, Kleerebezem R, van Loosdrecht MCM, van den Brand TPH. 2011. Selective sludge removal in a segregated aerobic granular biomass system as a strategy to control PAO-GAO competition at high temperatures. Water Research, 45(11):3291-9.

Winkler MKH, Kleerebezem R, Kuenen JG, Yang J, van Loosdrecht MCM. 2011. Segregation of biomass in cyclic anaerobic/aerobic granular sludge allows the enrichment of Anaerobic Ammonium Oxidizing Bacteria at low temperatures. Environmental Science and Technology, 45(17):7330-7.